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ApeCoin (APE) is the primary utility asset for the Yuga Labs ecosystem and the native gas token for ApeChain. Originally governed by a DAO, the project has evolved into a dedicated infrastructure layer for Web3 gaming and culture. ApeChain operates as a Layer 3 network built on Arbitrum Orbit, leveraging Ethereum’s security while offering the low fees and high throughput required for gaming applications.
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The ecosystem has undergone a fundamental restructuring to focus on product delivery and cross-chain utility:
ApeChain Mainnet Launch (October 2024): The launch of the ApeChain mainnet transitioned APE from a purely governance token to a network gas token. The chain introduces "Native Yield," allowing users to earn automatic yield on APE, ETH, and stablecoins (USD) bridged to the network without staking or locking assets.
Governance Overhaul (ApeCo): In June 2025, the community passed AIP-596 with over 99% approval, authorizing the dissolution of the ApeCoin DAO. Governance and treasury management are transitioning to ApeCo, a dedicated entity led by Yuga Labs, aimed at streamlining operations and removing the inefficiencies of the previous DAO structure.
Project R.A.I.D. (Cross-Chain Expansion): In late 2025, ApeCoin initiated "Project R.A.I.D," expanding the token's presence to non-EVM chains. APE is now natively integrated on Solana and BNB Chain, allowing for seamless transfers and usage across major blockchain ecosystems.
The APE token serves multiple functions across its supported networks:
Network Gas: APE is used to pay for all transaction fees on ApeChain.
Native Yield: Holders on ApeChain earn passive yield generated from the underlying protocol revenue and staking mechanisms.
Ecosystem Currency: APE remains the primary currency for Yuga Labs’ "Otherside" metaverse and is accepted by various third-party games and NFT marketplaces.
Token Standard: ERC-20 (Ethereum), Native (ApeChain), SPL (Solana)
Total Supply: Fixed cap of 1 Billion APE
Network Architecture: Layer 3 (Arbitrum Orbit Stack)
Governance Model: ApeCo (Centralized Stewardship transition from DAO)
Learn All About DAOs: How Decentralized Autonomous Organizations Work